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Clade
Subclade Archosauria: Avemetatarsalia to Neornithes 
Taxon Feedback
Taxon Dromaeosauridae 
Nominal Author Matthew and Brown 1922 
2° Nominal Author Russell 1969 
Taxon Status active
inactive
Comments The active definition is a first-order revision of the first definition in Sereno (1998), using species as specifiers. Following Padian et al. (1999), Dromaeosaurus albertensis is used as an internal specifier rather than Velociraptor mongoliensis to ensure inclusion of the eponymous genus. Velociraptor mongoliensis is better known but not appreciably more nested within the clade than Dromaeosaurus albertensis. The only coelurosaurs that have been placed by quantitative analysis as sister taxon to dromaeosaurids besides troodontids are ornithomimosaurs and birds. Nested specifiers from these two clades are cited as additional external specifiers to maintain the content of Dromaeosauridae under alternative phylogenetic arrangements. 
Potential Synonomy  
Active Phylogenetic Definition Feedback
Active Definition The most inclusive clade containing Dromaeosaurus albertensis Matthew and Brown 1922 but not Troodon formosus Leidy 1856, Ornithomimus edmontonicus Sternberg 1933, Passer domesticus (Linnaeus 1758). 
Shorthand > Dromaeosaurus albertensis but not Troodon formosus, Ornithomimus edmontonicus, Passer domesticus. 
Definitional Author Sereno 1998 
Definition Status original
textual substitution
first-order revision
second-order revision
Definition Type node
stem
Node-Stem Triplet yes
no
Other Triplet Taxa Deinonychosauria, Troodontidae 
Specifiers
Specifier A Dromaeosaurus albertensis 
Specifier(s) B  
Specifier(s) C Troodon formosus, Ornithomimus edmontonicus, Passer domesticus 
Qualifiers
+Taxon  
-Taxon  
Datum  
Taxonomic Content stem (entirely extinct)
crown (extant-bounded)
mixed (extant/extinct-bounded)
Publication Year 1998 
Inactive Taxon Status Feedback
Rejection Criteria junior synonym
historically polyphyletic
incomplete definition
no definition
apomorphy-based
redundant
other 
Critique  
 
Current Age Range Feedback
Earliest Record Hauterivian-Barremian (approx. 135 Ma) 
Latest Record Maastrichtian (66 Ma) 
Range (My) 69 
Basis for Range The earliest record is Utahraptor ostrommaysi from the Cedar Mountain Formation of Utah; the latest records are dromaeosaurids from the Maastrichtian of western North America, Asia and possibly southern Europe (Norell and Makovicky 2004). Dates are rounded to the nearest million, the earliest from the midpoint of the two listed stages and the latest from the end of the stage (Gradstein et al. 2004). 
Definitional History #1
Definition 1 All deinonychosaurs closer to Velociraptor than to Troodon. 
Author Sereno 1998:65 
Type node  stem 
Specifiers Dromaeosaurus, Velociraptor 
Qualifiers  
Definitional History #2
Definition 2 The most recent common ancestor of Dromaeosaurus and Velociraptor and all of its descendants. 
Author Padian et al. 1999:75 
Type node  stem 
Specifiers Dromaeosaurus, Velociraptor 
Qualifiers  
Definitional History #3
Definition 3 All taxa phylogenetically bracketed by Dromaeosaurus and Velociraptor. 
Author Senter et al. 2004:20 
Type node  stem 
Specifiers Dromaeosaurus, Velociraptor 
Qualifiers  
Definitional History #4
Definition 4 All the descendants of the most recent common ancestor of Microraptor zhaoianus, Sinornithosaurus millenii, and Velociraptor mongoliensis. 
Author Norell and Makovicky 2004:196 
Type node  stem 
Specifiers Microraptor zhaoianus, Sinornithosaurus millenii, Velociraptor mongoliensis 
Qualifiers