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Clade
Subclade Archosauria: Avemetatarsalia to Neornithes 

Taxon Feedback
Taxon Allosauroidea 
Nominal Author Marsh 1878 
2° Nominal Author Currie and Zhao 1993 
Taxon Status active
inactive
Comments

Currie and Zhao (1993:2079) and, independently, Sereno et al. (1994) were first to use the taxon Allosauroidea. Sereno (1997) identified Allosauroidea as stem-based, but did not provide a definition. Sereno (1998) gave a stem-based definition that includes closely related taxa rather than just allosaurids and sinraptorids. There are a number of well preserved forms, including genera such as Yangchuanosaurus, Monolophosaurus, and Cryolophosaurus, with close, yet, uncertain, affinities to Allosaurus and Sinraptor. In this case, a stem-based taxon is more useful to allow reference to the clade as a whole. If spinosaurids or some other tetanurine clade is repositioned within Neotetanurae and is more closely related to Allosaurus fragilis, then they would also be considered allosauroids.

Padian and Hutchinson (1997:6, fig. 1) provided a node-based definition for Allosauroidea (mislabeled in their figure as “Allosauridae” ), basing the definition on Allosaurus and Sinraptor. In place of Allosauroidea as used by Sereno (1997, 1998), Padian et al. (1999) and Holtz et al. (2004) used a stem-based Carnosauria.

Sereno and others have rejected the use of Carnosauria, as it has a long history as a traditional polyphyletic repository for a disparate array of large-bodied theropods (see Carnosauria). Gauthier (1986:9) tried to resuscitate Carnosauria as a clade, but he did not provide a phylogenetic definition and listed large-bodied taxa that now are recognized as having disparate relationships (including abelisaurids, allosaurids, and tyrannosaurids). Furthermore, because relationships among allosauroids (sensu Sereno) are poorly known, there is often no demonstrable difference in the taxonomic content of Allosauroidea and Carnosauria (sensu Padian et al. 1998). In a recent review (Holtz et al. 2004), for example, Allosauroidea is listed in a taxonomy within Carnosauria, but the supporting phylogeny shows a basal polytomy with no resolution of Allosauroide.

Given these considerations, it may be best to avoid retooling the taxon Carnosauria out of respect for its historical usage. Allosauroidea is best conceived as a stem-based taxon, the definition derived from a first-order revision of Sereno (1998).

 
Potential Synonomy Carnosauria, Allosauria 

Active Phylogenetic Definition Feedback
Active Definition The most inclusive clade containing Allosaurus fragilis Marsh 1877 but not Passer domesticus (Linnaeus 1758). 
Shorthand > Allosaurus fragilis but not Passer domesticus
Definitional Author Sereno 2005 
Definition Status original
textual substitution
first-order revision
second-order revision
Definition Type node
stem
Node-Stem Triplet yes
no
Other Triplet Taxa Neotetanurae, Coelurosauria 
Specifiers
Specifier A Allosaurus fragilis 
Specifier(s) B  
Specifier(s) C Passer domesticus 
Qualifiers
+Taxon  
-Taxon  
Datum  
Taxonomic Content stem (entirely extinct)
crown (extant-bounded)
mixed (extant/extinct-bounded)
Publication Year 1998 

Inactive Taxon Status Feedback
Rejection Criteria junior synonym
historically polyphyletic
incomplete definition
no definition
apomorphy-based
redundant
other 
Critique  
 

Current Age Range Feedback
Earliest Record Pliensbachian (190 Ma) 
Latest Record Tithonian (146 Ma) 
Range (My) 44 
Basis for Range The earliest record is Pliensbachian (Early Jurassic), the earliest age of an allosauroid (Cryolophosaurus), given that Allosauridae is a stem-based taxon. The latest record is Tithonian, based on Allosaurus fragilis (Holtz et al. 2004). Dates are rounded to the nearest million, the earliest from the beginning of the stage and the latest from the end of the stage (Gradstein et al. 2004). 

Definitional History #1
Definition 1 Allosaurus and Sinraptor and all descendants of their most recent common ancestor. 
Author Padian and Hutchinson 1997:7 
Type node  stem 
Specifiers Allosaurus, Sinraptor 
Qualifiers  

Definitional History #2
Definition 2 All neotetanurans closer to Allosaurus than to Neornithes. 
Author Sereno 1998:64 
Type node  stem 
Specifiers Allosaurus, Neornithes 
Qualifiers  

Definitional History #3
Definition 3 Allosaurus fragilis, Sinraptor dongi, their most recent common ancestor, and all of its descendants. 
Author Holtz et al. 2004:100 
Type node  stem 
Specifiers Allosaurus fragilis, Sinraptor dongi 
Qualifiers